Irreducible complexity is the proposal that some functional biological systems contain several interacting parts such that removal of a necessary part destroys the system's basic present function.
Does the dependence of a biological system upon multiple interacting components provide evidence concerning how that system originated?
Some biological systems cease to perform their normal function when an essential component is removed. Intelligent Design advocate Michael Behe used the term "irreducible complexity" to focus attention on systems whose present function depends upon several coordinated parts.
The concept concerns functional dependency in an existing system. A crucial distinction must be made between demonstrating that a modern system requires several parts and demonstrating that no evolutionary pathway to that system is possible.
This distinction lies at the center of the debate. Present-day dependency is experimentally testable. Historical claims about whether simpler ancestors could perform different functions require additional evidence.
Biologists can experimentally identify components required for particular present functions. Many molecular systems also contain optional, redundant, replaceable, or lineage-specific components, so dependency must be evaluated for each system rather than assumed.
Evolutionary critics of irreducible-complexity arguments propose that complex systems can arise through changes of function, recruitment of existing components, duplication and divergence, scaffolding, redundancy followed by loss, and other historical pathways.
For ancient systems, complete step-by-step histories are often unavailable. The existence of homologous components and evidence for possible intermediate functions can support evolutionary models without necessarily identifying every historical step.
The number of essential components varies by organism, system, environment, and the particular function being tested.
Irreducible complexity is directly relevant to Intelligent Design because it asks whether coordinated functional systems display characteristics that are difficult to produce through selectable incremental changes. The argument is strongest when it addresses actual evolutionary pathways rather than merely noting that a modern system has necessary parts.
Irreducible complexity identifies a legitimate question about the origin of integrated function, but present-day part dependency alone does not settle the historical question. Competing explanations must be compared with the biological evidence.